THE PROVISIONAL HYPOTHESIS OF PANGENESIS
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Every one would wish to explain to himself, even in an imperfect manner, how it is possible for a character possessed by some remote ancestor suddenly to reappear in the offspring; how the effects of increased or decreased use of a limb can be transmitted to the child; how the male sexual element can act not solely on the ovules, but occasionally on the mother-form; how a hybrid can be produced by the union of the cellular tissue of two plants independently of the organs of generation; how a limb can be reproduced on the exact line of amputation, with neither too much nor too little added; how the same organism may be produced by such widely different processes as budding and true seminal generation; and, lastly, how, of two allied forms, one passes in the course of its development through the most complex metamorphoses, and the other does not do so, though when mature both are alike in every detail of structure. I am aware that my view is merely a provisional hypothesis or speculation; but, until a better one be advanced, it will serve to bring together a multitude of facts which are at present left disconnected by any efficient cause. As Whewell, the historian of the inductive sciences, remarks, “Hypotheses may often be of service to science when they involve a certain portion of incompleteness, and even of error.” Under this point of view I venture to advance the hypothesis of pangenesis, which implies that every separate part of the whole organization reproduces itself. So that ovules, spermatozoa, and pollen-grains—the fertilized egg or seed, as well as buds—include and consist of a multitude of germs thrown off from each separate part or unit.